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"Father Randolph Holton was dead. Not literally, of course. But in his heart, he knew he could not live on...not after everything he had learned. Sitting at his dark oak desk in the corner of the dimly lit, cozy study, the white-haired, frail Catholic priest--even frailer of late--dipped an old quill into a container of jet-black ink and began to write..."
John Holton's father, a Catholic priest named Randolph, has been missing for two weeks. Unable to go to the police for help, since doing so would tarnish his father's reputation as a celibate priest, John takes it upon himself to search for his father.
Over the course of Lifestream, players take on the roles of both John in the present and his father, Randolph, in the past. While John searches for answers, players see the troubling events that led to Randolph's disappearance firsthand.